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Vander Wall, S.B. and Joyner, J.W. (1998) Secondary dispersal by the wind of winged pine seeds across the ground surface. The American Midland Naturalist Journal, 139, 365-373.
http://dx.doi.org/10.1674/0003-0031(1998)139[0365:SDBTWO]2.0.CO;2

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