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Agnèse, J.F., Berrebi, P., Lévêque, C. and Guégan, J.F. (1990) Two Lineages, Diploid & Tetraploid, Demonstrated in African Species Barbus (Osteichthyes, Cyprinidae). Aquatic Living Resources, 3, 305-311.
https://doi.org/10.1051/alr:1990031

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