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O’Toole, B. (2002) Phylogeny of the Species of the Superfamily Echeneoidea (Perciformes: Carangoidei: Echeneidae, Rachycentridae, and Coryphaenidae), with an Interpretation of Echeneid Hitchhiking Behaviour. Canadian Journal of Zoology, 80, 596-623.
https://doi.org/10.1139/z02-031

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