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Hatton, J.C. and Smart, N.O.E. (1984) The Effects of Long Term Exclusion of Large Herbivores on Soil Nutrient Status in Murchison Falls National Park, Uganda. African Journal of Ecology, 22, 23-30.
http://dx.doi.org/10.1111/j.1365-2028.1984.tb00670.x

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