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Golubtsov, A.S. and Krysanov, E.Y. (1993) Karyological Study of Some Cyprinid Species from Ethiopia. The Ploidy Differences between Large and Small Barbus of Africa. Journal of Fish Biology, 42, 445-455.
https://doi.org/10.1111/j.1095-8649.1993.tb00347.x

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